Evolution and Amplifying Feedback

Self-reinforcing feedback cycles organise the evolution of life on Earth. Here, I describe several types of evolutionary feedback cycles.
The evolution of abilities within a species
Evolution co-evolves capabilities:
- Genes
- Physical capacities, like strength and speed
- Senses such as sight and hearing
- Social behaviours like communication, and
- Toolmaking
In early human ancestors, for example, genetic changes affecting hand bones and brain development reinforced one another. As grip strength and precision improved, individuals could manipulate stones more effectively; as tool use became more important, natural selection favoured enhanced brains with planning ability and fine motor control. Over time, these traits co-evolved, each amplifying the adaptive value of the others.
(The untold story of our remarkable hands and how they made us human: New Scientist: 17 Feb 2026)
Evolution of the species proceeds through a self-reinforcing feedback process driven by natural selection and natural variation.
| Natural selection: Survival of the better adapted | ||
| The better-adapted next generation | The better-adapted breeding population | |
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| Natural variation | |
Diagram: This feedback cycle amplifies improved adaptation in (1) the breeding population and (2) the next generation.
In this amplifying feedback cycle, driven by natural selection and natural variation:
- The entire population is subject to natural selection. Those with advantageous traits tend to survive and form a better-adapted breeding population.
- The breeding population tends to produce a better-adapted next generation by passing on (1) their advantageous traits and (2) natural variations, e.g., genetic mutations. Then the cycle repeats.
Both causal links in this cycle are only tendencies:
- The breeding population will sometimes not be better adapted. For example, a natural disaster could decimate the population and, by chance, leave inferior breeders. When this happens, this cycle is not dominant.
- The next generation will sometimes not be better adapted. For example, the offspring may not inherit an adaptive trait, or a mutation is often not beneficial.
Once an adaptive trait appears, it takes many generations or intense selective pressure for it to spread widely through the population. While natural selection is a constant pressure in this cycle, genetic mutation occurs randomly and intermittently.
Despite this being a slow, intermittent process, it has dominated over vast timescales. It has generated complex features like eyes and the extraordinary diversity of species on Earth, each with finely tuned abilities and intricate relationships with other species, the web of life.
The co-evolution of predators & their prey
Another feedback cycle that has contributed to the evolution of life on Earth is the co-evolution of predator and prey abilities, such as those of cheetahs and impala. Over many generations, this cycle can progressively increase the abilities of both populations.
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| Increased CHEETAH abilities. | Increased IMPALA abilities | |
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Diagram: The feedback cycle amplifying abilities in both predators and prey
In this symmetric amplifying feedback cycle:
- An increase in the abilities of the cheetah population exerts a selective pressure on the impala population, e.g., faster cheetahs mean the impala need to become faster, which tends to
- increase the abilities of the impala population, exerting a selective pressure on the cheetah population, which tends to
- increase the abilities of the cheetah population, and the cycle repeats.
Cooperative evolution: microbiome
Cooperation also plays a critical role in evolution. For example, the symbiotic relationships between:
- Bees and flowering plants
- Coral and algae
- Cleaner fish and sharks
- Humans and their gut microbes.
Consider the co-evolution of humans and their gut microbes. Although humans and their gut microbes are distinct species, they form an integrated evolutionary unit, as they depend on each other for survival. The microbes pass to the next human generation through natural childbirth and breastfeeding. Humans benefit from the gut microbiome in many ways, but for simplicity, I will focus only on the microbes that aid human digestion. The amplifying feedback cycle described above for the evolution of abilities within a single species organises the co-evolution of humans and the gut microbes.
| Natural selection: Survival of the better adapted | ||
| The better-adapted next generation | The better-adapted breeding population | |
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| Natural variation | |
Diagram: This feedback cycle amplifies improved adaptation in (1) the breeding population and (2) the next generation.
In this situation, the “better adapted” includes:
- For humans, the ability to host the microbes by supplying them with nutrition and a stable environment, and
- For microbes, the ability to assist humans in digesting a wide range of foods. This ability would have been crucial during food shortages, which were often severe for early humans.
Specialisation in symbiotic evolution
In this symbiotic evolution between humans and their gut microbes, there was:
- the initial cooperative interaction between them,
- mutual benefit, which increased the reproductive success of both the humans and the microbes.
- modification of the environment for the microbes as the human gut became a honed niche for the microbes, and
- locked in cooperation, through the specialisation of both the humans and microbes.
The organisation of this specialisation is an example of a complementary system, another amplifying feedback cycle.
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| Microbes increase their ability to support human digestion of certain foods. | Humans reduce their ability to digest these foods in the absence of microbes and become increasingly dependent on the microbes. | |
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Diagram: The feedback cycle amplifying the ability of microbes to digest some foods and the human loss of this ability
This amplifying feedback cycle is a complementary relationship. In this cycle:
- The microbes tend to increase their ability to digest some foods.
- Humans tend to reduce their ability to digest these foods.
Mutually reinforcing amplifying feedback cycles.
Here we have a group of amplifying feedback cycles, all mutually supporting one another in the evolution of life on Earth:
- The evolution within a species,
- The co-evolution of predators and prey,
- The co-evolution of symbiotic species, and
- The evolution of specialised cooperation between species.
Here, as in many other systems, amplifying feedback cycles interact and reinforce one another. They sometimes produce extraordinary outcomes, such as the intricate web of life on our planet. At other times, they organise destruction; for example, global warming now threatens this very web of life in which humans have flourished.
The introduction to my counselling pages includes:
- Links to the other counselling pages, which include pages describing how this approach relates to other counselling practices and theories, are at the top of the introduction page
- References for all the counselling pages are at the end of the introduction page.
Loaded: 19 Dec 2025; Updated 2 July 2026.